arXiv:2608.05158v1 Announce Type: cross Abstract: In biological evolution, unconstrained mutation can lead to catastrophic outcomes: organisms may evolve enhanced capabilities while losing essential functions for survival. Nature's solution is \textit{developmental constraints}, where core regulatory genes remain anchored while peripheral genes adapt freely. We observe that current self-evolution algorithms for large language models lack analogous constraints. They optimize purely for capability, implicitly assuming safety will be preserved. Our experiments reveal this assumption to be dangerously wrong: models can \textit{misevolve} into powerful yet dangerous entities. Inspired by how Hox genes anchor body structure across $500$ million years of evolution, we propose \textbf{Circuit-Anchored Evolution (CAE)}. Using mechanistic interpretability, we identify a tiny \textit{safety circuit}, comprising less than $2$\% of model features, that causally mediates safety behaviors. We anchor ...